General: Deciduous shrub, loosely branched, 1-3 m tall; stems more or less erect, unarmed, covered with round, yellow, crystalline glands; bark brownish.
Leaves: Alternate, mapleleaf-shaped, 3-22 cm long, 3-24 cm wide, 5- to 7-lobed, the lobes cleft almost half their length, sparsely hairy to glabrous except for yellow glands; stalks from shorter to much longer than the blades, sparsely hairy.
Flowers: Inflorescence of numerous flowers in a 10-30 cm long, ascending to erect raceme; flower stalks 5-12 mm long, jointed, subtended by 1-3 small bracteoles, the lower ones often leaflike; petals white, 0.5-0.8 mm long, fan-shaped with an oblong basal claw; hypanthium widely flared and deeply saucer-shaped, 1.2-1.5 mm long; calyces brownish-purple or greenish, rarely nearly white, the lobes narrowly egg-shaped to oblong-lanceolate, 3-4 mm long; styles glabrous or hairy, about equaling the stamens and petals.
Fruits: Berries, nearly round, blue-black with a whitish bloom, 0.8-1.2 cm long, glandular.
Moist woodlands, forests, streambanks, shorelines, thickets and avalanche tracks in the lowland to the subalpine zones; frequent on the Queen Charlotte Islands, Vancouver Island and the adjacent mainland, mostly west of the Coast-Cascade Mountains; N to S AK and S to CA.
A shade-tolerant/intolerant, submontane to subalpine, Western North American deciduous shrub distributed more in the Pacific than in the Cordilleran region. Occurs in hypermaritime to maritime cool mesothermal climates on very moist to wet, nitrogen-rich soils; its occurrence increases with increasing precipitation and decreases with increasing elevation and continentality. Scattered in semiopen forests on water-receiving (floodplain and stream-edge) sites. Usually associated with Alnus rubra, Oplopanax horridus, and Rubus spectabilis. A nitrophytic species characteristic of Moder and Mull humus forms.
Recommended citation: Author, Date. Page title. In Klinkenberg, Brian. (Editor) 2019. E-Flora BC:
Electronic Atlas of the Plants of British Columbia [eflora.bc.ca]. Lab for
Advanced Spatial Analysis, Department of Geography, University of British
Columbia, Vancouver. [Accessed:
08/12/2019 9:03:24 PM
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