General: Plants perennial, loosely mat-forming, from a branched caudex.
Stems: Stems prostrate. Scapes solitary, sparsely to densely hairy with long, soft, shaggy hairs, 2-10 cm tall.
Leaves: Leaves in multiple dense basal rosettes, strap-shaped to oblong, margins ciliate, surfaces usually grey soft-hairy with simple hairs (rarely glabrescent), 3-15 mm.
Flowers: Inflorescences of 2-8 flowers; involucral bracts lanceolate to ovate. Flowers short-stalked; corollas salverform, white to pink-tinged with a yellow throat (throat turning pinkish with age), 4-8 mm wide; calyces broadly campanulate to subglobose, sometimes slightly keeled, long-hairy, lobes ovate-lanceolate 2-5 (8) mm; pedicels erect, 0.5-1.5 mm long, shorter than the calyces. Flowering Jun-Aug.
Fruits: Capsules globose or subglobose, about equalling the calyx, 5-valvate.
Source: The Vascular Flora of British Columbia, draft 2014. Author: Jamie Fenneman
Key to the Species and Taxonomic Notes
Key to Androsace
1a. Plants perennial, mat-forming, from a branched caudex; leaves grey soft-hairy, in multiple rosettes; flowers 4-8 mm wide, white with a yellow throat; pedicels shorter than the calyces………………………,,,..…………………..…….A. chamaejasme
1b. Plants annual or biennial, not mat-forming, from a taproot; leaves glabrescent to hairy with straight hairs but not grey soft-hairy, in a single basal rosette; flowers 1-3 mm wide, white; pedicels much longer than the calyces…………………………...2
2a. Involucral bracts broadly lanceolate to oblong or ovate, leaf-like; corollas essentially included within the calyces; calyces not keeled; calyx lobes narrowly ovate, about equal to the tube………………,,,……….………….………...……..A. occidentalis
2b. Involucral bracts linear to lanceolate, not leaf-like; corollas slightly exceeding the calyces; calyces keeled; calyx lobes more
or less triangular, shorter than the tube…………….………...……………………………………………..…A. septentrionalis
Source: Vascular Flora of British Columbia, draft version 2014
Author: Jamie Fenneman
Habitat / Range
Mesic to dry, calcareous meadows, gravelly areas, rocky slopes, heath, and tundra in the subalpine and alpine zones. Rare in n and se BC (Rocky Mts.); AK east to NU, south to NM; Eurasia.
Source: The Vascular Flora of British Columbia, draft 2014. Author: Jamie Fenneman
Additional Notes
This distinctive, showy species is closely associated with calcareous environments and coarse, well-drained soils. Our plants represent the North American component of a circumpolar complex; the remaining taxa in this complex occur in Eurasia. North American and northern Eurasian populations of A. chamaejasme have traditionally been attributed to ssp. lehmanniana (Spreng.) Hult.; however, as ssp. lehmanniana should properly apply only to European populations (at least populations from the Caucasus), northern and North American plants are better recognized as ssp. andersonii (Elven 2013). The distribution of this species, both in North America and Eurasia, is characterized by numerous widely disjunct populations.
Source: The Vascular Flora of British Columbia, Draft Version February 2014
Author: Jamie Fenneman
Family Information
Primulaceae:
Annual, biennial, or perennial forbs; scapose; from fibrous roots, short rhizome, or branched caudex, occasionally mat-forming or cespitose. Scapes ascending to erect, solitary or several per plant, unbranched. Leaves all basal, often rosulate, simple, unlobed, entire to toothed, subsessile to stalked, glabrous to pubescent, sometime glandular. Inflorescence a terminal, bracteate umbel, or flowers solitary. Flowers radially symmetric, 5-merous, bisexual, stalked; corolla campanulate or tubular to salviform, lobes sometimes sharply reflexed, usually whitish or pinkish to deep magenta, 4- to 5-lobed; calyces 4- to 5-lobed; stamens 5; filaments distinct or partially connate; ovary superior; style 1. Fruits 5-chambered dehiscent capsules, valvate to circumcissile. Genera 20, species ca. 600 (4 genera, 15 spp. in B.C.). Widespread in arctic, temperate, and subtropical regions of the northern hemisphere; disjunct in equatorial regions (e.g., e Africa, Indonesia) and in the southern hemisphere (e.g., s South America).
Non-scapose genera that were formerly included in Primulaceae ( Anagallis, Glaux, Lysimachia, Trientalis) have now been moved to Myrsinaceae (Källersjö et al. 2000, Trift et al. 2002, Cholewa and Kelso 2009). Many species in this family have a pronounced association with calcareous environments and are subsequently rather locally distributed within the province. Primulaceae contains a number of very showy species, and several genera have become important ornamentals in the horticultural trade (e.g., Primula, Dodecatheon).
Key to the Genera of Primulaceae
1a. Corolla lobes >2 times as long as the tube, sharply reflexed; stamens far exserted; filaments often at least partially connate (distinct in some species) .........................................................................Dodecatheon
1b. Corolla lobes less than twice as long as the tube, not reflexed; stamens included; filaments never connate…………………….........................2
2a. Calyces keeled (at least on the tube); plants densely cespitose (cushion-like) or mat-forming perennials with clusters of leaves at the bases of the scapes (rarely taprooted biennials), corollas rose-pink (fading to lavender, rarely to whitish).…...............................................Douglasia
2b. Calyces not keeled or only weakly keeled in fruit; plants taprooted annuals or perennials from fibrous roots or short rhizomes, or if mat-forming perennials (i.e., Androsace chamaejasme), then corollas white or pink-tinged with a yellow or pink throat..……………………….……………………………………………………………………………………...................................3
3a. Corollas constricted at the throat, magenta to lavender (rarely white), usually >7 mm across; plants never densely grey-hairy… ………………………………………………………...…………………………………………………..................................................Primula
3b. Corollas not constricted at the throat, white (sometimes fading pinkish in A. chamaejasme), usually <5 mm across (if >5 mm, then plants densely grey-hairy)…….……….......................................................……Androsace
Source: The Vascular Flora of British Columbia, draft 2014. Author: Jamie Fenneman
Genus Information
Annual or biennial, taprooted herbs or perennial, mat-forming herbs from a branched caudex. Scapes erect, solitary or several per plant, often elongating with age. Leaves obscurely stalked, hairy to sometimes glabrescent. Inflorescences terminal, bracteates, umbellate; branches often elongating with age. Flowers short- to long-stalked; corollas tubular to salverform, not constricted at the throat, white, sometimes with a yellow or pinkish centre, 5-lobed; calyces hemispheric to campanulate, 5-lobed; stamens included; ovary superior. Capsules globose or subglobose, about equalling the calyx, 5-valvate. About 100 spp. (3 spp. in B.C.); North America, Mexico, Eurasia.
This genus is centred in Eurasia, with few representatives in the New World. It is apparently paraphyletic without the inclusion of species that are currently placed in Douglasia (Schneeweiss et al. 2004, Boucher et al. 2012, Roquet et al. 2013), but that genus is retained as distinct here due to its unique morphology; see the account of Douglasia for more information. The genus is composed of two deeply divergent clades (Boucher et al. 2012, Roquet et al. 2013), although the morphological variation within each of these clades exceeds that which exists between them, making it difficult to identify any morphological characteristics that consistently separate the clades. A. occidentalis and A. septentrionalis (as well as the genus Douglasia) represent a European-American clade, while A. chamaejasme represents a Central Asian clade, with the divergence between these clades estimated at ca. 34 million years. This ancient divergence suggests that the two clades may warrant recognition as separate genera or, at least, subgenera.
Source: The Vascular Flora of British Columbia, draft 2014. Author: Jamie Fenneman
