Paired thorns on scapular region; a series of continuous thorns from nuchal region onto tail (Ref. 559). Tail long, its length equal to or longer than disc length (Ref. 559). Snout length 1/2 or more of distance from snout tip to 5th gill opening (Ref. 559). Ventral side of disk white, with grayish margin (Ref. 559).